brown to black when calling). If the female is receptive, she leaves the territory and approaches the calling male. The pair then moves to oviposition sites (e.g., a rock crevice or leaves). Clutch size is about 2–12 eggs measuring 3.5 mm (0.14 in) in diameter.[3]

The males tend the eggs until they hatch after about 21 days, and then carries the tadpoles to pools with slow flow rates, sometimes wet leaf litter when suitable pools are not available. Males generally avoid pools with the predators Anablepsoides hartii or Macrobrachium carcinus and may carry the tadpoles for long distance in order to find a suitable pool. It takes about 56 days for the tadpoles to reach metamorphosis.[3] While males may carry the tadpoles for up to four days, this seems not incur significant costs in terms of reduced feeding, fat loss, or locomotion (jump length and number).[2]

Distribution, habitat, and conservation :

The frog is found within the Northern and the Central Ranges. They are found along densely shaded, narrow, shallow, slow-flowing, clear water streams in undisturbed montane and moist forests. The frog is listed on the IUCN Red List as "vulnerable", being locally threatened by pollution, habitat loss and habitat degradation. Although it has a small geographic range, the frog is locally abundant,[1] and is not present on the CITES Endangered Species List.