Clams
Giant Clams, both the Hippopus Clams and the Tridacna Clams, are beautiful, hardy, grow rapidly, and require little care!
Caring For Tridacnid Clams
Giant Clam Care
courtesy to : www.animal-world.com Authors: Elizabeth M. Lukan / Clarice Brough, CFS.
Caring For Tridacnid Clams
Giant Clam Care
Giant Clam Care - Clam Facts
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Reef Aquarium SetupAquarium ParametersLighting Tips
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Giant Clams for Sale:What to Look for When Purchasing Giant Clams
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Placing Giant Clams In the Reef Tank
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Giant Clam CareFeeding Giant ClamsNutrients for Giant Clams
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Giant Clam Hitchhikers:
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Giant Clam Predators and Pests:
All of the above is true when speaking of a clam that is already well adapted to your aquarium. As with all pets (in your tank or otherwise), they must be taken care of properly in order to actually live up to those claims. Once established a giant clam makes an awesome addition to a reef aquarium. Though like all reef inhabitants, it will need good water quality and proper lighting, it will require little else in the way of care.
Giant clams can give you a number of years of enjoyment, however the initial adjustment to your tank can be a considerably difficult time. The differences in lighting, pH, temperature, salinity, water current, and more can be drastic to the new clam. It's important to be as well prepared as you possibly can for the care of your clam before actually purchasing one
Try to purchase farm raised giant clams instead of collecting them from the wild. Tridacnid clams have been exterminated in many areas because of over harvesting, collected for food and shells.
The mariculture of tridacnid clams was begun to re-stock areas where the clams had been eliminated, and to provide a farm raised source of clams for food. A portion of the farm raised clams now goes to the aquarium trade. Profits from the demand of aquarists has raised interest in producing colorful varieties of all the species. Ask your fish stores and online sellers for farm raised clams.
Reef Aquarium Setup
Reef aquarium parameters:
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Temperature: Mid to Upper 70° F. (mid 20° C.). Do not let the aquarium exceed 84° F.
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Salinity / Specific Gravity: 1.024. Salinity is also important, too high or low a salinity can cause the death of a clam. Try to keep specific gravity between 1.023 and 1.025.
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pH: 8.3. Do not let the aquarium exceed a pH above 8.4.
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Hardness: Maintain a dkh of 7.9.
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Water Movement: For most giant clams low to moderate currents will be tolerated.
Reef Aquarium lighting :
Giant Clams will do well under moderate to relatively high lighting intensities. They don't particularly like intense lighting nor will they tolerate sudden increases in intensity. Take great care if attempting to acclimate them to intense lighting such as metal halides. T. gigas can be adapted to metal halide lighting, but this should be done over time as a gradual process.
When determining the lighting needs of your clam, a few basic things to consider are; placing it in the aquarium, its mantle color, what species of clam is it, and its needs as it ages:
Lighting tips for keeping giant clams:
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Placing giant clams: With brighter lights, you can place your clams lower in the tank. With lower intensity lights, you will be forced to place them near the surface.
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Giant clam species - lighting needs: Light requirements are different for the different species of clams:
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The Crocea Clam T. crocea and Maxima Clam T. maxima need the most light..
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These are followed by the Gigas Clam T. gigas, Derasa Clam T. derasa, and Squarmosa Clam T. squamosa.
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The least amount of light is needed by the Hippopus Clam H. hippopus.
  Most clams need a lot of blue light in the bulb's spectrum. The Derasa Clam T. derasa needs less blue light than Gigas Clam T. gigas and Maxima Clam T. maxima.
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Clam mantle color - lighting needs: Clams with brown mantles do not require as much light as those with blue mantles, usually, and are considered easier to keep in the home aquarium lighting wise. Clams with blue mantles are usually found in shallow-water. The blue pigment acts as a light filter and so they require substantial quantities of light. A good rule to stick to is, the more colorful the clam, the greater quality and quantity of light it will need.
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Giant clam ages - lighting needs:: As the clam grows, its mantle will thicken and the number of zooxanthellae will increase (with the deeper lying zooxanthellae receiving less light) and it will require more and more light. Smaller clams, those less than 1.56 inch / 4.0 cm, require less light to maintain their optimum growth while the larger clams require more light. Also, juvenile clams will adapt to lighting variables more readily than adult clams.
Changes in lighting / environment - effects on giant clams:
When you take a clam from the sea and place it in the aquarium, the difference in the spectral composition of the light can have drastic impact on the clam's food supply. A clam at sea receiving the full light spectrum transferred to a tank with the same light intensity, only with blue wavelengths dominating, is receiving the same amount of light but some wavelengths are missing and others are in greater quantities.
Depending on the clam's adaptation, some of the zooxanthellae's assimilation pigments may cease to function and pigments that would use the new wavelengths aren't even present. Basically in this situation, the new lighting is incorrect for the clam even though the intensity of the illumination is the same as before. The clam can adapt to the new spectral composition, slowly, but the time needed for this to occur may be too long.
This leaves the clam weakened, with less resistance to defend against predators, diseases, etc. Smaller clams require less light.
Giant Clams for Sale:
What to look for when purchasing Tridacnid clams - what to know about giant clams for sale:
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Watch for gaping Clams: Recently imported or transported clams usually exhibit a behavior called gaping. A gaping clam will appear as follows: shell fully open, mantle poorly extended, and inhalant siphon widely stretched. This eventually passes. Gaping will continue if the clam is kept under insufficient lighting, is damaged, or unhealthy. The mantle will begin to pull inwards, shriveling and tearing between the siphons. A healthy clam's inhalant siphon can open wide sometimes, but gaping leaves a very wide opening. The clam will stay like this for as long as the clam is unhealthy.
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Examine the mantle: The clams mantle should be colorful everywhere with no clear or white areas. Colorless areas may be the result of poor lighting, predators, or disease. A clam will quickly recover from poor lighting once conditions are improved.NOTE: It is normal for T. gigas to have clear areas near the center of their mantle. Also check for rips and tears in the mantle. A healthy mantle should be extended over the edge of the shell and not pulled inwards. NOTE: It is normal for H. Hippopus' mantles to not extend over the shell.
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Check the clam's reactions. A healthy clam should react to external stimulus by forcefully closing its shell. Newly imported and transported clams tend to react more slowly, but will improve as they regain their strength.
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Check the clam's byssus gland: The byssus gland should be undamaged. You should not see any torn or loose tissue hanging from the bottom of the clam. Some byssal strands may be visible, but no solid tissue hanging loose. Byssal gland damage isn't always visible, however. The clam may appear fine for a couple of weeks and then die suddenly for seemingly no reason. On the good side, byssal gland damage isn't always fatal. According to The Reef Aquarium Volume One, Delbeek and Sprung have collected and purchased damaged clams with little loss.
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Removing a giant clam: If the clam is attached to substrate, please take care when detaching. Lift the shell gently and insert a sharp knife, razor, or scissors and cut the threads as close as possible to the substrate. Do not cut close to the shell. You could cut into the extended byssal gland. If the clam is attached to a small rock, pebbles, etc., just leave it alone. The Crocea Clam T. crocea and Maxima Clam T. maxima are very sensitive about being handled like this and are best left attached if possible.
Farm raised giant clams: Hatchery (culture/farm) systems for giant clams are setup basically the same. Baby clams only live a short time in tanks. Once the clams reach 0.75 to 1.25 inches (20 to 30 millimeters) they are moved to ocean nurseries. They spend the rest of their growing time, until they reach about 1.5 inches (4 centimeters) or more, in the ocean. This is true for majority of the clams, regardless of their species, that are marketed to the hobby. So, even the farm raised clams are from the sea!
A hint for clam sellers: A good way to display clam stock would be to place them on small cups or pots filled with crushed coral.
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This way the clam will only attach to the gravel. This makes it very easy to remove the clam when sold and with much less stress on the clam and its byssal threads and gland. There is also less stress on a seller who has to dismantle their display to detach the clam while the customer stands around waiting.
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Rinse the gravel routinely to prevent worms from moving in.
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The pots will also help keep the clams upright and positioned properly under the light source.
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Dealers can also hang a mirror above the aquarium (at a 45 degree angle). The customer will then see the true colors of the clams without having to contort themselves.
Placing Giant Clams In the Reef Tank :
When putting your clam in your tank there are a few placement considerations:
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Clam movement: Once placed, clams generally cannot move around on their own. This is why where you put them in your tank is of vital importance. Clams placed on hard surfaces (rocks) will not be able to upright themselves or shift their positions. Clams on sand can manage to upright themselves or shift positions with small movements. Don't expect your clam to move any great distances. Juveniles clams settle permanently, only using their foot to travel short distances. Generally the larger the clam, the less moving around you should expect, which basically means your clam's place in your tank and its happiness there is really up to you.
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Closing clam shells: Clams can close their shells with enough force to expel a surprising amount of water out of their siphons. If you place your clam near the top of your tank, this water may be expelled out of the tank or up into your lighting system. Clam's may also accidentally trap small, slow moving fish that rest on their mantle (for example, mandarins, gobies, hawkfish, or blennies).
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Surface - flat or inclined: Find as flat a surface as possible and place the byssal opening flat on the substrate with the mantle facing directly up. Horizontal surfaces are best for clam placement. If you insist on putting your clam on an incline, make sure the byssal opening is on the lower portion of the substrate. If the incline is too great, the clam will not receive enough light, so do not place it on steep substrate.
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Substrate types: A clam may fall over several times before it firmly attaches to the substrate. A good idea is to put some small rocks (crushed coral, large pieces of substrate, etc.) around the clam to help it stay upright. These small rocks will not get in the way of the opening and closing of the shell. Your clam will attach within a few days to a week. Substrate is not related to attachment speed. Once your clam is attached, you can remove the rocks, unless the clam has used them in its attachment. Do not place your clam between large rocks, inside small holes, or up against the tank wall, or you may prevent them from opening fully.
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The Crocea Clam T. crocea and Maxima Clam T. maxima are found in rocky habitats so it is best to place them on rocks. .
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The Squarmosa Clam T. squamosa, Derasa Clam T. derasa, Gigas Clam T. gigas, and Hippopus Clam H. hippopus are best placed on sandy substrates.
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Lighting requirements: Remember the lighting requirements of the clam you are putting in your tank. Clams with colorful mantles need a great deal of light. Clams with brown mantles, not as much light. If your clam has a brown mantle, place it nearer the bottom of your tank, or shield it (an overhang would work well for this) from strong lighting if you have to.
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Giant clam position:
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Don't leave a clam that has fallen over, upright it as soon as possible.
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Inhalant siphon: Place the clam so the inhalant siphon (which lies above the byssal opening) is on the lowest portion of the slope. As the clam grows it will place greater strain on its byssal gland, so if the gland is on the upper portion of the slope, the weight of the clam could gradually pull the gland out. Adult Hippopus sp. tend to sit more on their hinge than on the byssal opening.
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Mantle: Giant clams should be placed so that the majority of their mantle is facing upwards. Juveniles will attach themselves to rocks with byssus threads just as other tridacnid clams do.
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Currents: GIant clams do not like strong currents, especially the Crocea Clam T. crocea. Do not place them where they would receive strong, direct water currents. Too much current will cause your clam not to open. They do need water flow to bring nutrients to them, just not too strong.
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Tank mates: Keep your clam as far away from any aggressive coral or anemone as is possible in your tank. If sections of the mantle are pulled away or shriveled on the same side as a coral or anemone, it is probably irritating the clam and move the coral, anemone, or clam as soon as possible. Do not hesitate, or it will die quickly.
Giant Clam Care :
With proper lighting and careful attention, giant clams require little else in the way of care. It is important to make sure they are not being irritated, not being fed upon by other organisms, and good water quality must be maintained. Giant Clams can be kept in a reef environment with live rock and coral substrates. The number one cause of a giant clam's demise is usually water quality. A high pH and high temperatures can cause problems.
Feeding Giant Clams:
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Giant clams receive the majority of their nutrition from their zooxanthellae, whether additional feeding is required is still debated. Some hobbyists believe that tridacnids should be fed, going on the assumption that they are filter feeders like other clams.
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According to The Reef Aquarium Volume One, when Delbeek and Sprung attempted to feed the clams, they closed forcefully and expelled the food. They may accept a dilute suspension of live phytoplankton or a yeast. However, the effort required to feed these items is not worth it in the opinions of Delbeek and Sprung. Delbeek and Sprung noted that for many years, Tridacnid clams have been grown successfully in both culture systems and home aquaria without any supplemental feedings. They noted that bacteria, organic and inorganic compounds are always present in the water of closed systems (like our tanks), and these may be consumed or absorbed by the clams.
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Of opposing opinion, Albert Thiel noted that in some cases supplemental feeding may be necessary. What those "cases" are was not mentioned. Mr. Thiel further notes that small foods should be used and that clams do not feed on large chunks of food. Use good quality food like shrimp or scallop meat run through a blender.
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For more detailed information about the nutrition and feeding requirements of clams see: Giant Clams - What Do Clams Eat?
Basic Nutrients for giant clams:
Basic nutrients in the aquarium that these giant clams need are calcium, strontium, iodine, magnesium, and possibly a minute amount nitrate.
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Calcium: Calcium is the main building block for clams and should be present in the water at levels of at least 280 mg/L for growth to occur. More rapid, natural growth is seen when calcium is in the range of 400-480 mg/L.
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Strontium: Strontium is incorporated in the shell along with calcium and should also be provided for optimum growth.
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Iodine: The addition of iodine to the aquarium will also enhance growth and color in giant clams.
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Magnesium: Magnesium aids in maintaining proper calcium levels and in the formation of skeletal material in clams
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Nitrate: They require some nitrogen for proper growth. Nitrate can be added if levels are extremely low, but be careful as nitrates should never exceed 2 mg/L.
Giant Clam Hitchhikers:
What might be hitching a ride on a giant clam:
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Clam shell with encrusting organisms: Clam's sometimes arrive with growths of encrusting organisms on their shells, especially the Maxima Clam T. maxima and Squamosa Clam T. squamosa. Check these growths very carefully. Dead or necrotic areas may foul your tank.
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Parasitic snails: Various parasitic snails can be imported with your clam. They are carnivorous boring snails and that description should be enough for you to want them out of your tank and as far away from your clams as possible. Look for small rice grain-sized, cream colored spots near the base or hidden within the flutes (the grooves and indentations in the shell) of the shell, or, at night, along the upper edge of the shell. If the clam is attached to a rock, check by lifting the clam a short distance off the rock and look underneath. You are looking for small (0.08-0.2 inch / 2-5 mm long) snails. Remove all of these snails.
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Quarantine giant Clams: If you have the facilities, quarantining your clam until you are sure all the snails are removed is a good idea. Also check for the egg masses of these snails and remove them as well. They are small, jelly-like masses on the shell. Don't confuse the jelly-like mass some clams excrete around their byssus opening for these egg masses .Keep looking for these snails. Just because you think you got them all when you added the clan to your tank doesn't mean you really did!
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Safe shrimps and crabs: Symbiotic shrimp from the family Palaemondidae (Anchistus, Conchodytes and Paranchistus) or small crabs such as Pea Crabs of the family Pinnotheridae may be visible through the inhalant siphon in larger clams. These animals live inside the clam and do not harm it, although what they eat and what they do for the clam is unknown.
Giant Clam Predators and Pests:
In the wild, small tridacnid clams are heavily preyed upon. Many species of fish (triggerfish, large wrasses, puffers, etc.), crabs, lobsters, shrimp, polychaetes (Bristleworms, Fireworms, etc.), octopi, and snails prey on clams. Even burrowing sponges! Since most of the clams available to aquarists are juveniles, hobbyists should be extra cautious about both predators and environmental factors that can wreak havoc with a clam.
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Fish Predators: Certain wrasse species (Family Labridae) are bad tank mates. Species like the Twin Spot Wrasse Coris aygulaand Bird Wrasse Gomphosus varius have been known to attack and devour juvenile clams in the aquarium. Sometimes the clam is eaten from above or knocked over and eaten through the soft and unprotected byssal opening. Any large wrasse species should be watched closely when introduced into a tank with giant clams. Other fish can also irritate clams. Fish that are constantly grazing like as Centropyge sp. (Pygmy Angelfish),Ctenochaetus sp. (Chevron Tang, etc.) and Acanthurus sp. (Achilles Tang, etc.) Tangs, and Ecsenius sp.(Blennies) will occasionally nip at a clam in passing. Sometimes small pieces of tissue are removed but the problem is that the clam is now irritated and it will expand less and less. It may eventually expand so little that it won't receive enough light and will slowly die.
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Crab Predators: Large crabs will eat clams and usually shortly after they are placed in the tank, before they can attach to the substrate. They attack the clam through the byssal opening, but larger crabs just crack the shells open.
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Shrimp Predators: Certain species of shrimp can also prey on clams. Large shrimp such as Marble Shrimp Saron marmoratus and Buffalo Shrimp Saron sp. will attack clams at night. The common Cleaner Shrimp Lysmata amboinensis has been known to attack clams. It's rare, but it does happen, especially when the clam is injured and the shrimp hungry.
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Snail Predators: Parasitic snails are sometimes imported with tridacnid clams as mentioned earlier. Examine new clams closely and remove any snails or eggs right away.
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Environmental factors: A number of environmental factors can also irritate giant clams, creating potential problems:
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Algae:Algae is another problem for clams. If algae begins to grow over the lip of the shell, the mantle may become irritated and it will not expand as much.
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Macroalgae:Macroalgae like Caulerpa can irritate the clam from underneath if allowed to grow under the byssal opening. If this happens, the clam will produce large amounts of mucus from below that surrounds the base.
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Clam Mucus:Producing large amounts of mucus is a normal means of protection for the clam against algae, stinging corals, or predators. The mucus is thick, clear, and often contains brown patches (brown jelly). The effect of noxious by-products of soft corals (example: Xenia spp.) can also cause clams to produce large amounts of clear mucus. The mucus can quickly clog prefilters
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Aiptasia anemones:If Aiptasia are allowed to grow on tridacnid clams, they can reach underneath the mantle and sting the clam. This will result in the mantle pulling away and the clam will eventually die.
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Clam worms:Polychaete worms such as the larger Nereis sp. and Eunice sp. can prey upon tridacnids. They are usually active at night and feed on the clam from below, through the byssal opening or by boring a hole through the shell.
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Air bubbles:Air bubbles can be a problem too. They can become trapped inside the clam and cause the clam's demise.
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For more information on giant clam predators and pests see: Tridacnid Clams: Friends, Enemies & Ailments
Aquarium Invertebrates: A Look at the Giant Clam Tridacna maxima
courtesy to: Pomacanthus Publications, Inc.
With their desirability in mind, if you can find a good specimen these clams can be relatively easy to care for in a well-run reef aquarium. However, they do have particular lighting requirements, and are by no means bulletproof when it comes to keeping them long-term. So, this article will cover their basic biology, how to identify them, and how to best care for them in aquaria.
There are several species of clam belonging to the family Tridacnidae, which are best known as the tridacnids or giant clams. Of these, one of the most attractive species is Tridacna maxima, which is also one of the most commonly offered species available to hobbyists. I say most attractive because they can come in a wide range of colors, which can be arranged in a variety of unusual patterns, with many specimens being striped, sprinkled, spotted, blotched, marbled, etc. The colors themselves also range from black and white, with essentially everything else in between being seen on some specimen or another. In fact, I'd say it's harder to find a maxima that's unattractive than to find one that is.
Basic Information
To get started, maxima is the most widely distributed species of the tridacnids. They're found in the Red Sea and from East Africa all the way across the Indo-Pacific to Polynesia. They also live as far north as southern Japan, and as far south as the Great Barrier Reef (Rosewater 1965). Maximas can be found in high numbers around many reef areas where waters are relatively shallow and clear, with the majority living at depths less than about 25 feet. Some can be found living as deep as about 50 feet, but their abundance drops off dramatically below about 25 feet, with these deeper-living clams occurring mostly as solitary individuals (Jaubert 1977).
Maxima range
Regardless of their depth of occurrence, essentially all of them are found living on limestone substrates, on top of living corals, or on coral rubble. Supposedly they're occasionally found on sandy bottoms (Pasaribu 1988), but after doing a lot of diving around Japan and Indonesia I have yet to see this. Regardless, on hard bottoms maximas can chemically bore a shallow indentation into the substrate that the bottom of their shells fits into, and they strongly affix themselves in place using a tough structure called a byssus. So, they typically stay in one spot for life, with the bottom third or half of the shell kept out of sight in their burrow. Conversely, on coral rubble bottoms they simply bury themselves amongst the coral chunks and attach to something solid with their byssus if they can. Again, usually only part of the shell rises above the substrate. The odd thing is that they won't do this in aquariums, though. It seems that if they don't start making a burrow while they're relatively tiny, they won't do it at all. So, don't expect a specimen to dig into your live rock. Regardless, they almost always attach to the substrate using their byssus anyway.
Aside from that, the most notable thing to point out here is that, like all the other members of the family, maximas harbor large populations of zooxanthellae. These single-celled photosynthetic algae live in the tissues of a host clam primarily within a specialized system of tubes that permeate the fleshy, colorful, mantle tissue that extends from the top of the shell, and when given enough light, they can make far more food than they need for themselves. The extra food (in the form of carbon and energy-packed glucose) is then given to the clam host, which is the same thing that occurs within most reef-dwelling corals.
Under optimal conditions, these zooxanthellae are constantly multiplying within a tridacnid, and some of these live algal cells can be digested by specialized amoeboid cells within the host, too. So, a host clam can rely on its zooxanthellae for more than just sugar, and is considered to be a "farmer" to some degree since it can consume these surplus zooxanthellae grown inside its body.
In addition, all tridacnids can also absorb a variety of nutrients directly from seawater. Their fleshy mantle is covered by a specialized tissue that can very effectively take in dissolved nutrients like ammonia, nitrate, and phosphates. So, here they have a third means of nutrient acquisition, with one more to go.
The last way they cover their nutritional needs is through filter-feeding. All tridacnids can eat fine particulate matter strained from surrounding waters by their specialized gills, which not only work to exchange carbon dioxide and oxygen, but can also act as sieves that can collect such particles. A tridacnid, like most other clams, pumps water into its body chamber, where it flows over the finely-branched gills and then flows out the other end of the body chamber, minus some particulates. These collected bits are can include phytoplankton, zooplankton, and detritus, meaning they can make use of a broad range of things.
Identification :
When it comes to identification, once you know what to look for maxima is usually pretty easy to distinguish from all other tridacnids with the exception of T. crocea. So, I'll go over the basic features used to ID them, and then give you some tips on how to differentiate them from croceas, too.
When it comes to shells, they're almost always grayish-white when clean. However, one of the interesting things about the shells of this species is that sometimes they may be tinted with light yellow or pinkish-orange. Rarely, the shell may also be completely yellow. It's almost always strongly elongated in form, being much longer than it is tall, and some maximas are very thin from side to side while others are quite fat. Deformed shells are not particularly uncommon either, as maximas sometimes live in very crowded groups and/or partially burrowed into coral rock preventing them from producing a normally-shaped shell. Regardless, at its top each half of the shell typically has four or five smoothly-curved and inter-digitating projections that are symmetrical to those on the other, allowing the them to close together tightly. However, there are occasional individuals that have more elongated and even pointed tooth-like projections that don't inter-digitate as smoothly with those on the opposite side.
Some species of tridacnids have petal or shelf-like structures on their shells, which are called scutes, and maxima is one of them. In fact, their shells are typically covered by numerous tightly-spaced but thin scutes, which run in rows from the bottom to the top of the shell. However, when maximas partially burrow into the substrate, many of these scutes are either not formed in the first place, or are broken/eroded away in the process. So, maxima shells oftentimes have no scutes on the bottom portion, while numerous scutes are still present on the rest. Still, there are occasional individuals that have none at all for some reason, while aquacultured specimens that are not permitted to burrow typically retain most or all of their scutes.
Also note that it's possible for a maxima's shell to reach almost 16 inches in length, but that's the largest ever reported (Kinch 2002). Thus, you shouldn't expect any given specimen you purchase to get so big. In fact, McMichael (1974) did a survey of several hundred maximas in the wild and reported that only 3% were larger than 9 inches and the largest specimen found in the whole survey was only 9.8 inches. So, that record holding 16-inch specimen was quite an anomaly.
When it comes to the soft parts, maximas typically extend their zooxanthellae-packed mantle tissue well beyond the upper edges of the shell. In fact, it's typically extended to the point that it completely obscures the shell from view when looking down on one. The mantle can also come in such a wide range of colors and patterns that there really is no standard color, although blue is the most common. As noted, the patterns covering it may also be striped, sprinkled, spotted, blotched, marbled, etc. and quite fancy, which is why various specimens are often called things like teardrop maximas, striped maximas, super maximas, or even ultra maximas, etc.
Still, the only patterns that are relatively consistent in how they look are that of the teardrop and striped varieties. Teardrop maximas may vary significantly in color, but they tend to have the same sort of pattern covering their mantle, being covered in teardrop-shaped splotches, while striped maximas tend to have a dark, solid background color with thin radiating stripes of blue, yellow, or white. Other than that, the mantle has rows of simple, closely-spaced, dark eyes near the outer edge and sometimes has numerous eye-tipped tubercles/protrusions on its upper surface, too. The large mouth-like opening in it (called the inhalent siphon) is also ringed with numerous simple, small tentacles that usually lack anything more than very fine branches.
The dark spots on these maxima's mantles are simple eyes.
A typical teardrop maxima.
A typical striped maxima.
Now, as I said above, maximas are indeed easily confused with croceas because both species have relatively large and often brightly-colored mantles with small tentacles around their inhalent siphons. The vast majority of maximas has elongated shells with lots of scutes, while almost all croceas have shorter, taller shells that lack scutes or only have a few small ones. But, there are exceptions, which lead to this confusion. A typical crocea's shell lacks scutes and is far less elongated than the shell of a typical maxima, but there are individuals of each species that are in between. Croceas can be rather elongated at times, and may actually have a lot of scutes, while some maximas may have rather short shells and lack scutes. So, I'll give you some additional pointers for trying to figure out which is which in case it isn't clear who is who.
Some maximas are not elongated (L), while some croceas are (R). This crocea even has a few rudimentary scutes, which are often larger and more numerous on aquacultured specimens.
First, a maxima's shell usually has larger, much more pronounced waves or folds than that of a crocea, as crocea's shells are typically relatively smooth. A maxima's shell sometimes has very sharply-pointed, almost triangular projections at the shell's upper edge, but crocea's are always more rounded and never sharp. Maximas can reach significantly larger sizes than croceas, as the record-holding crocea was only 6 inches long. So, anything larger than about 5 inches in length is almost certainly a maxima. There is no such thing as a teardrop crocea or a striped crocea. Croceas may have some stripes on them at times, but I've never seen one that had a solid background color with thin radiating stripes on top, or the characteristic droplets of a teardrop. And lastly, the tentacles around the inhalent siphon of a maxima are typically simple and un-branched, while those of a crocea are usually finely branched at their tips.
The tentacles surrounding maxima's inhalent siphon are typically simpler than those of crocea.
So, there is no straightforward single way to always ID both species correctly, but by looking at a combination of these features you can usually figure out just about any of them. I will admit though, over the years I've come across a handful of specimens that have been quite difficult to differentiate. At such times most folks just throw up their hands and declare that a hard-to-ID specimen in a hybrid between the two species, but after doing a lot of searching, reading, and asking clam farmers questions I'm still far from convinced that these two species can/do hybridize. That's a topic for another day, though.
Aquarium Care :
When it comes to caring for maximas, water quality requirements are typical for reef aquariums in general. Basically, if you're successfully keeping corals alive and well, then your water quality is good enough for a maxima. On the other hand, if you're having problems maintaining excellent water quality - don't fool with any species of tridacnid.
When it comes to water motion, tridacnids live in reef and near-reef environments, and are regularly exposed to strong currents and wave activity. This is especially so for maximas, which often live right at the crest of a reef where waves break hardest. Thus, they are no strangers to strong, surging and turbulent water motion. However, in aquariums the flow tends to be quite linear and constant, as a pump outlet might blast water in one particular spot day and night at about the same volume per minute, and rarely creates any real surge or turbulence. So, you need to think about this when it comes to the placement of a maxima (or any other tridacnid) in an aquarium.
It's okay to expose maximas to a low velocity surge, or to turbulent flow, but putting them in a position where a pump constantly hits them with a strong, non-stop linear current is not recommended. Basically, any sort of current that causes the mantle to fold upwards too much, or over onto itself all the time is bad, as is any current that makes a specimen chronically retract its mantle. Thus, you can put one anywhere you like with respect to current, as long as it doesn't bring on either of these reactions. I'll also add that while they're almost always found on hard substrates and rubble in the wild, placing them on such is highly recommended, but is not required. Placing a specimen on sand/gravel won't kill them, but they often move around a lot, trying to find something to attach their byssus to. Next is lighting, which not surprisingly is of critical importance.
Maximas live at relatively shallow depths where they receive relatively intense light, so fluorescent lighting will only suffice in shallow tanks, or if a specimen is placed on the rockwork near the water's surface in a deeper tank. I would try fitting as many bulbs into the canopy/fixture as possible at that, and mount the bulbs close to the water, and then place any specimens within a foot of the surface, preferably less. Some specimens may be able to get by at times with less light, or further down in deeper tanks, but I implore you to not take chances. Metal halide or comparable L.E.D. lighting is your best option.
I know that some people have gotten by with less, but when it comes down to it insufficient lighting is certainly one of the most common causes of losses. The problem is that corals are very simple organisms that have no real "guts" to speak of, while tridacnids have all the organs you'd expect to find in a higher animal. They've got stomachs, kidneys, gonads, gills, and even a heart. Thus, they are far more complex than you might think, and they use a lot of calories to keep everything running. So, it's a mistake to think that just because your lighting is bright enough to keep corals healthy and growing that they're necessarily bright enough to keep a maxima alive long-term.
To make matters worse, it can take a tridacnid months to slowly starve to the point of no return. So, everything can look fine for weeks on end, then a specimen may seem to just up and die for no apparent reason when it was really starving the whole time. Every maxima is genetically different at that, and long-term experience has proven that some individuals can get by with less while others need much more, even though they may be the same species and even the same size and color. To add, you cannot give a tridacnid too much light as long as a specimen is given time to adapt to intense lighting, so it's better to err on the bright side than the dim side. For more on this, refer to my article On Lighting for Tridacnid Clams in the March 2011 issue, and for even more than that see my book Giant Clams in the Sea and the Aquarium.
You can't overdo it when it comes to lighting, as many maximas are found in the intertidal zone where they're exposed to tropical sunlight that's as bright as it gets.
Lastly, there's the question of whether or not you need to feed a maxima in an aquarium. As covered, all tridacnids are filter-feeders, yet their zooxanthellae can cover a great deal of their nutritional needs, and they're able to absorb pretty much everything else they need directly from seawater. In fact, if provided with enough light, maximas of any size have no need to filter feed and can thrive in particulate-free water as long as there are enough dissolved nutrients present. Controlled experiments by Fitt & Trench (1981) proved that tridacnids can do without, and I kept them for years before anyone was talking about adding phytoplankton to aquaria, much less sold any in a bottle. You can get all the details in my article Tridacnid Clams (Usually) Don't Need to Be Fed in Aquaria in the July 2010 issue, but I'll give you some basic info on the subject anyway.
When you feed your fishes some amount of the food won't get eaten and becomes detrital particles, which maximas can filter out. Any uneaten food also releases nutrients into the water as it decomposes. Likewise, the food that is eaten by the fishes ends up becoming solid wastes that can also become detritus. However, even more importantly, fishes excrete dissolved substances that can be absorbed by a clam, too. For example, fishes give off dissolved ammonia as a waste product, but tridacnids can absorb it and use it as a source of nitrogen. Thus, when you feed your fishes, you're feeding your tridacnid(s), too.
So, the real question is whether or not there are enough fishes in your aquarium to support one or more tridacnids. While it's unlikely to happen, I suppose it is possible to have too low a fish load (or too high a tridacnid load depending on how you look at it) in an aquarium, which would mean that the amount of fish waste being produced would not be enough to support the needs of the clam(s). So, my advice is to refrain from taking any chances and use a quality phytoplankton product if you have any doubts. I have to say though, I imagine that very, very few hobbyists have problems due to nutrient levels that are too low since for most of us the fight is to prevent them from getting to high.
Anyway, I need to wrap things up, and unfortunately I'll going to end on a bad note. As gorgeous as they may be, most all experienced aquarists agree that maxima is the least hardy of the tridacnids. I've seen and heard of more losses of this single species than all the rest by far, and I think it's usually due to insufficient lighting. They are especially dependent on excellent water quality and intense lighting. So, if you don't have both, don't buy one of these.
I'll also add that despite their attractiveness, availability, and relatively low price, really small specimens are even more likely to pass away. In fact, I experienced so many losses of small maximas back in my selling days that I outright refused to order/sell them after a while, and I've heard the same from many other vendors, too. They didn't tolerate shipping and acclimation to aquarium life very well at all, and it was common for more to die in the first couple of weeks than to live. Stick with larger specimens, as in at least few inches long, and you'll have much better odds of success.
References :
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Fitt, W.K. and R.K. Trench. 1981. Spawning, development, and acquisition of zooxanthellae by Tridacna squamosa (Mollusca, Bivalvia). Biological Bulletin 161:213-235.
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Jaubert, J. 1977. Light, metabolism, and the distribution of Tridacna maxima in a South Pacific atoll: Takapoto (French Polynesia). Proceedings of the 3rd International Coral Reef Symposium 1:489-494.
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Kinch, J. 2002. Giant clams: their status and trade in Milne Bay Province, Papau New Guinea. TRAFFIC Bulletin19(2):1-9.
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McMichael, D.F. 1974. Growth rate, population size and mantle coloration of the small giant clam Tridacna maxima (Roding), at One Tree Island, Capricorn Group, Queensland. Proceedings of the Second International Coral Reef Symposium 1:241-254.
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Pasaribu, B.P. 1988. Status of giant clams in Indonesia. In: Copeland, J.W. and J.S. Lucas (eds.) Giant Clams in Asia and the Pacific. ACIAR Monograph Number 9, Canberra. 274pp.
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Rosewater, J. 1965. The family Tridacnidae in the Indo-Pacific. Indo-Pacific Mollusca 1:347-396.